| |
During the following discussion, keep in mind that we are talking specifically
about flowering plants-rather than the more primitive mosses, for example.
Although some of the processes are the same for both groups, it will save
us from having to add qualifiers such as "except for this or that group
of plants."
Let's begin with a summary of reproduction in animals.
In animals, meiosis results directly in the formation of gametes-the reproductive
cells, egg and sperm. The successful union of an egg and a sperm is called
fertilization, and the resulting
cell is called a zygote. The zygote
contains genetic material donated from both parents, and is genetically
distinct from either.
In plants, there are some intermediate
steps in the process. Rather than producing egg and sperm, plants use meiosis
to produce spores.
(Mitosis and meiosis are covered in detail in Section
C, Plant Growth.) Let's take a closer look.
In the anthers of a flower, certain diploid cells undergo meiosis, forming
haploid spores (called microspores-"little
spores"). These microspores divide by mitosis to form two-celled pollen,
consisting of a tube cell and
a generative cell.
If the pollen reaches a receptive stigma, the pollen germinates.
The
stigma has a sticky surface so that pollen can easily adhere to it. This
transfer of pollen from anther to stigma is known as pollination.
The tube cell germinates and produces a pollen
tube which grows down the style. The long, hollow pollen tube will
eventually penetrate the pistil's tissues in search of the egg. At the same
time, the generative cell divides by mitosis to produce two sperm. The two
sperm will travel down the pollen tube and enter the ovule, which contains
the egg.
What egg? The female gametes, or eggs, are formed in
the ovules, within the ovary in the flower. Here,
as above, a certain diploid cell undergoes meiosis. However, in this case,
the result is four haploid megaspores
("large spores"). Three of the megaspores die and disintegrate. The
remaining cell undergoes mitosis several times. One of these daughter
cells then functions as an egg, ready to be united with a sperm.
One sperm fuses with the egg to create a single diploid cell, the zygote.
This is fertilization.
The zygote cell then divides, growing into the multi-celled embryo.
The other sperm combines with another cell within the ovary-called the
polar nuclei-forming the endosperm. Recall that the endosperm will serve
as a temporary food supply that nourishes the developing embryo.
|
|